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Don,
The bypass capacitors in a divider chain don't usually have either
of their plates connected to ground. They still supply charge to the
photomultiplier tube.
The fact that some of that charge may end up back
on the other plate in no more relevant than the fact that current that
comes out of one terminal of a power supply goes into the other.
Would we
say that a power supply never supplies current just because the NET
current out it is zero?